- Cell Walls are found in plant, fungi, and bacteria cells.
Functions
- It maintains turgor pressure within the cell which gives plant cells their characteristic polyhedral shape.
- They serve as skeletons for both the cell and the organism.
- Protects against damage from abrasion and pathogens
- Mediates cell-cell interactions.
Composition
- They contain a fibrous element embedded in a non-fibrous gel-like matrix.
- Cellulose makes up the fibrous elements of the cell wall. They are organized into microfibrils that make the cell wall rigid and add tensile strength.
- Microfibrils are often organized in cross-hatching fashion, where the microfibrils of one layer are oriented perpendicular to those of adjacent layers.
- Cellulose is synthesized via cellulose synthase on the cell surface.
- The cell matrix consists of the non-fibrous parts which are synthesized within the cytoplasm
- Hemicellulose - branched polysaccharides that bind and cross-link microfibrils to form a resilient network
- Pectins - negatively charged glacturonic acid-containing polysaccharides. They form a hydrated gel that fills the space of the fibrous elements.
- There are proteins that facilitate dynamic activities
- Expansins - facilitates relaxation of the cell wall at certain sites.
- Kinases - transmit signals from cell-wall to cytoplasm.
Growth
- They arise from a thin cell plate which then matures by incorporate additional materials.
- Primary Cell Walls are the flexible walls of growing cells. Secondary Cell Walls are the thicker walls found in mature plant cells.
- The transition from primary to secondary occurs due to an increase in cellulose.
- They often incorporate lignin for additional structural support.
Links
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Extracellular Matrix and Extracellular Interactions - more on cell-cell interactions.