• Cell Walls are found in plant, fungi, and bacteria cells.

Functions

  • It maintains turgor pressure within the cell which gives plant cells their characteristic polyhedral shape.
  • They serve as skeletons for both the cell and the organism.
  • Protects against damage from abrasion and pathogens
  • Mediates cell-cell interactions.

Composition

  • They contain a fibrous element embedded in a non-fibrous gel-like matrix.
  • Cellulose makes up the fibrous elements of the cell wall. They are organized into microfibrils that make the cell wall rigid and add tensile strength.
    • Microfibrils are often organized in cross-hatching fashion, where the microfibrils of one layer are oriented perpendicular to those of adjacent layers.
    • Cellulose is synthesized via cellulose synthase on the cell surface.
  • The cell matrix consists of the non-fibrous parts which are synthesized within the cytoplasm
    • Hemicellulose - branched polysaccharides that bind and cross-link microfibrils to form a resilient network
    • Pectins - negatively charged glacturonic acid-containing polysaccharides. They form a hydrated gel that fills the space of the fibrous elements.
    • There are proteins that facilitate dynamic activities
      • Expansins - facilitates relaxation of the cell wall at certain sites.
      • Kinases - transmit signals from cell-wall to cytoplasm.

Growth

  • They arise from a thin cell plate which then matures by incorporate additional materials.
  • Primary Cell Walls are the flexible walls of growing cells. Secondary Cell Walls are the thicker walls found in mature plant cells.
    • The transition from primary to secondary occurs due to an increase in cellulose.
    • They often incorporate lignin for additional structural support.

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